The study says that the Ethiopians studies "had hemoglobin
concentrations that differed little from sea level
values. The similar findings of erythropoietin
concentration and oxygen saturation in the normal sea
level range were consistent. The findings would not be
remarkable if this were a sea level sample, but it is
a sample of people exposed to chronic, lifelong
hypobaric hypoxia. Ethiopians must have unique
adaptations of oxygen uptake or delivery that result
in the absence of an hypoxemic stimulus to increase
red blood cell production and hemoglobin concentration
despite their high-altitude residence."
They have sea-level concentrations of hemoglobin yet they've been at altitude for 40,000 years. The way I read this, their hemoglobin carry more oxygen than non-Ethiopians.
"The results of this study suggest that Ethiopian
high-altitude natives respond to hypobaric hypoxia
differently than Andean or Tibetan highlanders. This
Ethiopian sample resembles samples from the Tibetan
Plateau in exhibiting little or no elevation of
hemoglobin concentration in this altitude range, but
differs from Tibetans who have very low oxygen
saturation. It differs from Andean high-altitude
natives who exhibited both substantial elevation of
hemoglobin concentration and low oxygen saturation.
These findings suggest there are three patterns of
adaptation to high-altitude hypoxia among indigenous
populations (Table 1). Learning why the three
populations differ will require two lines of future
investigation. One is understanding the biological
mechanisms and the underlying genetics that allow
successful high-altitude adaptation with
quantitatively different suites of traits for oxygen
sensing, response, and delivery. The other is
understanding the evolutionary processes that produced
these patterns to explain how and why several
successful human adaptations to high altitude evolved."